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Bettany first gained popularity for appearing in the films Gangster No. 1 (2000), A Knight's Tale (2001) and A Beautiful Mind (2001). He was nominated for a BAFTA Award for playing Stephen Maturin in Master and Commander: The Far Side of the World (2003). Other films in which Bettany has appeared include Dogville (2003), Wimbledon (2004), The Da Vinci Code (2006), The Tourist (2010), Margin Call (2011), Legend (2015) and Solo: A Star Wars Story (2018). He made his directorial debut with the film Shelter (2014), which he also wrote and co-produced.

In television and theatre, Bettany has portrayed Ian Campbell, 11th Duke of Argyll in the series A Very British Scandal and Andy Warhol in the play The Collaboration in the West End, which later came to Broadway.