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Arnett was Emmy-nominated for his recurring role as Devon Banks in the NBC sitcom 30 Rock (2007–2013). He starred in the NBC sitcom Up All Night (2011–2012), the CBS sitcom The Millers (2013–2014), and the Netflix mystery series Murderville. He has also starred, co-created, and executive produced the Fox sitcom Running Wilde (2010–2011) and the Netflix comedy-drama series Flaked (2016–2017).

Arnett has acted in films such as Let's Go to Prison (2006), Blades of Glory (2007), Semi-Pro (2008), G-Force (2009), and Teenage Mutant Ninja Turtles (2014). Known for his deep baritone voice, Arnett also took roles in Danny Phantom (2004–2007), Ice Age: The Meltdown (2006), Ratatouille (2007), Horton Hears a Who! (2008), Monsters vs. Aliens (2009), Despicable Me (2010), The Nut Job film franchise (2014–2017), The Lego Movie franchise (2014–2019), Teen Titans Go! To the Movies (2018), Chip 'n Dale: Rescue Rangers (2022), and Twisted Metal (2023–).

Arnett has been Primetime Emmy Award-nominated for Arrested Development, 30 Rock, and BoJack Horseman. Since 2020, he has hosted the Fox reality series Lego Masters. Arnett began co-hosting the comedy podcast SmartLess (2020–) alongside Sean Hayes and Jason Bateman.