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Kilner's breakthrough role came in 1994, when he was cast as William Boone in the science fiction series Earth: Final Conflict. The series ran for five seasons, and Kilner's performance as Boone was praised by critics. He was nominated for a Saturn Award for Best Actor on Television in 1997.

In 1995, Kilner starred in the sitcom Almost Perfect, opposite Nancy Travis. The show was canceled after two seasons, but Kilner's performance was again praised by critics.

In 1999, Kilner starred in the family film Smart House. The film was a critical and commercial success, and Kilner's performance as Nick Starkey was well-received.

Kilner has continued to work in television and film since then. He has appeared in a number of other television series, including The Practice, Dollhouse, and The Good Wife. He has also appeared in a number of films, including Raising Helen, A Cinderella Story, and Home Alone 3.